Dendroecological studies of historical tree recruitment patterns suggest mixed‐severity fire effects are common in Douglas‐fir/western hemlock forests of the Pacific Northwest (PNW), USA, but empirical studies linking observed fire severity to tree regeneration response are needed to expand our understanding into the functional role of fire in this forest type.

Altered fire regimes can drive major and enduring compositional shifts or losses of forest ecosystems. In western North America, ponderosa pine and dry mixed‐conifer forest types appear increasingly vulnerable to uncharacteristically extensive, high‐severity wildfire.

Western U.S. wildfire area burned has increased dramatically over the last half‐century. How contemporary extent and severity of wildfires compare to the pre‐settlement patterns to which ecosystems are adapted is debated. We compared large wildfires in Pacific Northwest forests from 1984 to 2015 to modeled historic fire regimes.

The frequency, extent, and severity of wildfire strongly influence the structure and function of ecosystems. Mixed‐severity fire regimes are the most complex and least understood fire regimes, and variability of fire severity can occur at fine spatial and temporal scales, depending on previous disturbance history, topography, fuel continuity, vegetation type, and weather.

As wildfire activity increases in many regions of the world, it is imperative that we understand how key components of fire‐prone ecosystems respond to spatial variation in fire characteristics.

Prescribed fire is an active management tool used to address wildfire hazard and ecological concerns associated with fire exclusion and suppression over the past century.

Prescribed fire is an active management tool used to address wildfire hazard and ecological concerns associated with fire exclusion and suppression over the past century.

Wildfire ecosystems are thought to be self‐regulated through pattern–process interactions between ignition frequency and location, and patterns of burned and recovering vegetation. Yet, recent increases in the frequency of large wildfires call into question the application of self‐organization theory to landscape resilience.

Building resilience to natural disturbances is a key to managing forests for adaptation to climate change. To date, most climate adaptation guidance has focused on recommendations for frequent‐fire forests, leaving few published guidelines for forests that naturally experience infrequent, stand‐replacing wildfires.

A warming climate, fire exclusion, and land cover changes are altering the conditions that produced historical fire regimes and facilitating increased recent wildfire activity in the northwestern United States.